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  Home >>Zoology Dictionary >> Clavicle - Convergent Evolution

Clavicle - Bone of the pectoral girdle.

Cleavage - After fertilisation an egg cell undergoes many divisions to produce many small cells. The process is called cleavage.

Clitellum - The 'saddle' a thickened region of skin in earthworms and some other annelids. Concerned with reproduction, it is conspicuous only in mature animals.

Cloaca - Posterior opening of the body of most vertebrates into which open the gut, the excretory ducts and the reproductive system. Only the placental mammals have a separate anus.

Caidoblast - Stinging cel1 jellyfish and other Coelenterata
.
Conidoci - 'Trigger' of coelenterate stinging cell.

Coccyx - Rudimentary tail skeleton consisting of fused vertebrae.

Cochlea - That part of the ear which is concerned with the sorting out of the sounds received.

Cockroach - Insect of the order Dictyoptera.

Cocoon - A protective covering. Earthworms and many spiders wrap their eggs in cocoons. Many caterpillars spin siken cocoons around themselves as a protection when they are about to pupate.

Coelenterata - A phylum of animals which are radially symmetrical and whose body wall is composed of only two layers of cells ectoderm and endoderm separated by a layer of jelly like material the mesogloca. The body has a single cavity the coeknterol1 which has a single opening the mouth surrounded by a ring of tentacles. All coelenterates live an water and the majority are marine but a few, such as Hydra, live in fresh water.

Coelenterates are carnivorous animals, feeding on a variety of animals, feeding on a variety of animals which they catch by means of stinging cells found on the arms or tentacles. These stinging cells are called nematoblasts or cnidoblasts and each contains a. small capsule called a nematocyst. Inside the nematocyst there is a hollow thread that is inverted like the finger of a glove that has been pulled into the hand part. A passing animal perhaps a water flea in the case of Hydra or a small fish in the case of a seaanemone stimulates the tiny 'trigger' (cnidocil) of the stinging cell and the thread is shot out from its capsule. Some threads have poison barbs, others are coiled or sticky, and they all help to capture the prey.

The food is taken into the mouth and digestion begins. Some of the cells of the endoderm secrete enzymes that act upon the food and break it down: other cells possess whip like flagella that keep the liquid moving around. Digestion is completed within certain cells which engulf small particles just as Amoeba does. Most of the cells of the body and arms are called musculo epitheial cells because they possess muscular processes running through the mesogloea. These 'muscle tails' produce movement in the animals by contraction. The tails of the ectoderm cells run along the body while those of the endoderm run around the body. Contraction of the outer muscle tails shortens the body long and thin The arms and body bend by contraction of the outer muscle tails of one side only.

Many coelenterates possess two distinct forms in their life history a fixed, tubular polyp and a free swimming, disc shaped medusa but some groups have either polyp or medusa only. When both stages are present the polyp reproduces asexually by budding off mudusae and the medusae produce sex cells which fuse in pairs and develop into new polyps (See Alternation of Generations). The polyp stage often develops into large branching colonies by budding.

There are three classes within the phylum: (1) Hydrozoa coelenterates normally formingbranc,hed colonies and having both polyp and medusa (e.g. Obelia). There is, however, great variation in the life history. Hydra itself is very atypical of the group as it is solitary and has no medusoid stage.

(2) Scyphozoa coelenterates with an inconspicuous or absent polyp stage. The medusa may be very large. These are the typical jellyfishes.

(3) Actillozoa solitary and colonial forms with no medusae. The coelenteron contains vertical partitions (mesenteries). Corals have a calcareous sketeton secreted by the ectodrm and are frequently colonial. Sea anermones are usually solitary and never have a skeleton but they are nevertheless basically similar to corals and are included in the same class.

Coelom - Coelenterate animals like Hydra have a body wall with only two cell layers ectoderm and endoderm. Higher animals have another layer the mesoderm between these layers and. because they have three layers. they are called triploblastic animals. The mesoderm forms the muscles. reproductive organs, etc. In flatworms and other simple creatures it also forms a sort of packing tissue that separates the internal organs from the body wall andfrom each other. In more advanced animals, however, the mesoderm becomes split into two regions one surrounding the gut and other internal organs, the other lining the body wall. The fluid filled cavity between the two layers is the coelom. It is well developed in vertebrates, echinoderms, and many worms, but in arthropods and molluscs it is reduced and the main body cavity in these animals is really part of the blood system (a haemocoel). Animals with a coelom are said to be coelomate.

Coelomoduct - A duct of mesodermal origin connecting the coelom to the outside of the body e.g. the vertebrate oviduct. Eggs are shed into the coelom and then carried to the outside in the coelomoduct.

Coleoptera - Beetles. This order of Insecta is characterised by the possession of horny front wings (elytra) that meet in the mid line and normally cover all of the abdomen. All have biting jaws but they feed on a wide range of materials, some are fiercely carnivorous. Metamorphosis is complete, there being a larval stage very unlike the adult.

Collagen - A tough fibrous protein which binds various tissues together and also forms tendons and ligaments where high tensile strength is required.

Collembola - Order of primitive inects springtails.

Colon - Large intestine of vertebrates.

Comb Jellies - (See Ctenophora).

Commensalism - A state where two or more species of an animal live in fairly close association but without having much influence on each other. One species may benefit hut not both.

Commissure - Bundle of nerve fibres joining two parts of the central nervous system. E.g. circum oesophageal commissure joining the ganglia above and below the oesophagus of the earthworm.

Conditioned Reflex - The smell or taste of food makes a dog's saliva now. This is a normal reflex action involving no learning by the dog, but it can' be conditioned so that it occurs with a different stimulus. It a bell is sounded every time food is given the dog will associate the bell with food and soon the bell alone will cause
saliva to flow. This is the conditioned refex.

Condyle - Rounded projection of bone, especially on the skull, that fits into a socket on another bone to allow limited movement. The occipital condyles at the back of the skull articulate with the aflas vertebra and allow the head to be nodded.

Cone - Colour sensitive cell of the vertebrate retina.

Conjugation - A type of reproduction found in some protozoans such as Paramoecium. It involves paitial fusion of two individuals and the exchange of nuclear material Conjunctiva. Thin layer of tissue covering the front of the eye, and lining the eyelids of vertebrates.

Connective Tissue - This term is applied to various body tissues all of which bind cells and organs together. Connective tissue underlies the skin, surrounds nerves and muscles, joins bones and muscles to each other and often stores fat in its cells. Connective tissue cells are normally well spaced in a matrix of fluid in which there are numbers of collagen and other fibres. Tendons which join muscles to bones have to be very resistant to stretching and are made almost entirely of collagen, whereas other connective tissues may have very few collagen fibres.

Contractile Vacuole - Feature of many protozoans, especially fresh water species. There may be more than one in each organism and they gradually swell as they extract water from the surrounding protoplasm. Suddenly the vacuoles burst and force the water out of the body. They then begin to fill up again. This may be a means of discharging waste from the body but the main function appears to be the removal of water which continually passes into the body because of asmosis.

Conus Arteriosus - Muscular chamber of primitive vertebraie heart at beginning of ventral aorta.

Convergent Evolution - Hermann Melville, in his great sea story 'Moby Dick', wrote, To be short, then, a whale is a spouting fish with a horizontal tail. Whales do of course resemble fish. They are streamlined; they have a fluked tail, a pair of flippers as fore limbs, and they have warm blood and the females suckle their young with milk. Thus, despite all appearances, they are far more closely related to dogs, rabbits and Man himself than any fish.

One hundred and fifty million years ago another fish-like creature swam the seas. Though it became extirict 70 million years ago, fossils of its bones enable its, appearance to be reconstructed. But, far from being a fish, the preserved skeletons show beyon all doubt that the creature was a reptile. Because of its remarkable resemblance to a fish it was called ichthyosaur or fish lizard (Greek, ichthys, a fish; sauros, a lizard).The connecting link between whale, fish and ichthyosaur is; all have lived in the sea and during their evolution became adapted to an aquatic life. This evolutionary development, which often leads to apparent likenesses between stocks that have diverged a long way from an original ancestor, is called convergent evolution or simply convergence.

Convergence is a common phenomenon found in organisms living on land, in water and in the air. It provides powerful support for the theory of evolution by natural selection under the same selective pressures different groups will respond with apparently, similar adaptations. Care must be taken in classifying animals and plants: superficial resemblances between two organisms do not necessarily mean that they are closely related.

Cut off in the continent of Australia, the marsupials or pouched mammals have flourished for 150 million years. In most other parts of the world they have disappeared, for competition from the placental mammals proved too much.

Australia offers a wealth of environments and many different marsupials evolved, filling the niches.The similarity in appearance between the various marsupials and placental mammals elsewhere provides striking examples of convergence.

Thus Australia has its wolf the Tasmanina wolf the Tasmanian wolf, it has its native 'cat' (Dasyurus) and its native 'mouse' (Dasycercus). There are 'ant eaters' and there are 'stoths'. The flying phalanger is comparable with the flying squirrels and Australia even has its own 'marsupial mole.

Marsupials and placentals are all mammals and do possess a common ancestry in the not too remote past. Convergence can, however, produce similarity between completely unrelated creatures. Thus insects and birds are far removed from one another yet at a glance it is very difficult to tell the difference between a humming birds and a humming bird hawk moth in flight. Both are similar in size and both live off nectar in flowers and have converged in their hovering flight and their feeding procedures.

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