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Home >> Biology >> Morphology and Anatomy of Flowering Plants >> Origin and Activity of Cambium
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1. Origin and activity of cambium

Intrafascicular of fascicular cambium is present in dicot stem bundle in between the xylem and pholem of each vascular bundle. During secondary growth the parenchymatous cells of medullary rays become meristematic and form strips of interfascicular cambium from a complete ring of vascular cambium. The vascular cambium consists of two types of cells.

(a) Fusiform initials. These are spindle shaped elongated cells having brick like appearance. They produce secondary xylem elements and secondary pholem elements. These are arranged in vertical rows.

(b) Ray initials. These are arranged horizontally and produce pholem and xylem rays.
The cells of cambium ring divide by their tangential walls to form two daughter cells of which one remains meristematic while the other matures into secondary vascular tissue. The daughter cells formed on the outer side mature into secondary pholem and those produced towards inner side mature into secondary xylem and secondary continue division to form secondary xylem and secondary pholem.

The cambium is active towards inside than outside so secondary xylem increases more rapidly than secondary pholem pushing the cambium ring outwards. In the mean time some cells cambium divide by radial walls to increase circumference of cambium ring.

The ray initials cut off parenchymatous cells towards outside as well as inside to form radial bands of parenchymatous cells passing through secondary pholem and secondary xylem as secondary medullary rays.
Many of natural fibres such as jute, hemp, flax, etc. are bast fibres of secondary pholem. The primary pholem present outside the secondary pholem gets crushed. The secondary xylem forms the bulk of stem called wood. The primary xylem persists as conical projections on inner side of secondary xylem around pith.

Cambium Homogeneous Uniseriate
Cambium - Homogenous Uniseriate
Cambium - Homogenous Multiseriate
Cambium - Homogenous Multiseriate
Cambium - Heterogenous Multiseriate Rays
Cambium - Heterogenous Multiseriate Rays
Cambium - Square Upright Cells
Cambium - Square Upright Cells


Diagram of Cut Surface of a Dicotyledonous Stem Annual Rings
Diagram of Cut Surface of a Dicotyledonous Stem Annual Rings

Diagrammatic Representation of a Portion of Transverse Section of an Old Dicotyledonous Stem

Diagrammatic Representation of a Portion of Transverse Section of an Old Dicotyledonous Stem

1. Periderm
5. Spring Wood
2. Secondary Phloem
6. Autumn Wood
3. Modullary Ray
7. Pith

Annual rings. Activity of cambium is not uniform in plants growing in varying climate. In temperature climate, cambium becomes more active in spring while in winter it becomes less active.

The wood formed in spring is called spring wood and that formed in cold winter or dry summer is called autumn wood. These two woods appear as concentric rings known as annual rings. Each annual ring represents one years growth. The age of plant can be determined by counting the number of annual rings.

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